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makethumbs creates several smaller versions of images. If you are constrained for disk space, or want to keep the downloads small, you can reduce the compression setting. The default is 75%; JPEG can often be compressed down to around 40-50% before you start to see really obvious compression artifacts. It depends a lot on the type of image.
This option is most often used when your image galleries are hosted on an ISP with a small disk quota. You can usually be safe setting the compression down to 50-60% and shave a third size off your image sizes or so.
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